New Book available with Jean Raffaëlli - Nouveau livre disponible : La passion des Salamandres


After two published books : “Les Urodèles en voie de disparition” (2016, Edilivre, Paris, 120 p.) and “Les Salamandres crocodiles du genre Echinotriton en Asie du sud-est: de véritables fossiles vivants au bord de l'extinction” (2016, Edilivre, Paris, 112 p.); Pr. Axel Hernandez has decided to publish this year with Chimaira edition  a new comprehensive  study of the Primitive Salamandridae from Asia (Genera Echinotriton and Tylototriton). This new book is intended to summarize all the knowledge on each 25 species. This research is fed by the author’s field studies in Asia with chinese Amphibian specialists such as Pr. Hou Mian, Pr. Liang Fei, Pr. Ye Changyuang, Dr. Xie Feng, Dr. Jiang Jianping & Dr. Zhiyong Yuan (China, Thailand, Myanmar, Vietnam and Laos) since 2001; and improve our understanding of this marginal group of poorly known salamanders. The book includes : A preface made by one of the most famous amphibian specialist Professor Liang FEI (Chengdu Institute of Biology, Academy of Sciences, China), Acknowledgements, Travel notes from southern Asia, The Primitive Salamandridae of Asia (Natural History), Mass extinction of salamanders around the world, Phylogeny of the two genera with a new classification, Taxonomy, Biogeography, Ecological data for each 25 species known to this date, detailed pictures and maps, reproduction and captive husbandry for conservation purposes. This book is dedicaced to the world famous herpetologist and specialist of Salamanders, Jean RAFFAËLLI author of "Les Urodèles du monde" I & II (2007, 2013; Link: and co-author of the New Taxonomy of the entire Urodela group with the famous french scientific: Professor Alain DUBOIS (2009, 2012). His thoughts and knowledge illuminate and give life to this study.


The last decade saw numerous new taxa being described within these genera, emphasizing that our understanding the complexity within this group is still at an early stage. At the same time, urodeles are disappearing at an unprecedented pace, and it is to be hoped that the information provided here will help with conserving them in and ex situ. To this end, it was attempted to summarize all knowledges for each 25 species: Echinotriton maxiquadratus, Echinotriton chinhaiensis, Echinotriton andersoni, Tylototriton verrucosus, Tylototriton shanjing, Tylototriton pulcherrimus, Tylototriton anguliceps, Tylototriton podichthys, Tylototriton uyenoi, Tylototriton yangi, Tylototriton shanorum, Tylototriton himalayanus, Tylototriton kweichowensis, Tylototriton taliangensis, Tylototriton pseudoverrucosus, Supraspecies asperrimus, Tylototriton asperrimus, Tylototriton hainanensis, Tylototriton notialis, Tylototriton ziegleri, Supraspecies wenxianensis, Tylototriton wenxianensis, Tylototriton dabienicus, Tylototriton broadoridgus, Tylototriton lizhengchangi, Tylototriton liuyangensis, Tylototriton vietnamensis, Tylototriton panhai;  including morphological diagnoses and previous taxonomical studies, variation observed, distribution, habitat, ecology, ethology, and new data for individual species on their reproductive biology and captive breeding requirements that may prove useful for conservation breeding efforts, as well as information on their present conservation status, threats and conservation measures.


In this new book the author also applies the supraspecies concept suggested by Dubois & Raffaëlli (2009, 2012) and Raffaëlli (2013) in line with the ICZN Code. A supraspecies is a rank applicable to groups of closely related species below subgenus and above species. In Tylototriton, the rank of supraspecies is particularly useful to express in taxonomic terms divergences that have become apparent through molecular phylogenies. One could use terms like superspecies, species group or species complex, but none of these are of taxonomic relevance as per the Code. The concept of "epispecies" should be recognized by the Code and would be a good option to group some very closely related species within supraspecies like those I treat here as clades in the subgenera Tylototriton and Yaotriton.





The first complete book on Crocodile Salamanders with 415 pages, more than 600 pictures, a new classification, comprehensive maps and many more...


Découvrez la nouvelle Salamandre algire de l'Atlas (Salamandra algira atlantica) décrite récemment du Maroc. Elle mesure entre 16 et 20 cm de long et se caractérise par son aspect luisant, sa coloration originale, noire, tachetée de jaune, mais aussi de rouge !

 Hernandez and Escoriza, 2019, ZooKeys, 893 : 146. Holotype: MNCN 50499, by original designation. Type locality: "Jbel Tazekka, Taza Province, Middle Atlas Mountains, northern Morocco (34.15N, 4.00W) at 810 m a.s.l". / 



Tylototriton yangi

Tylototriton kweichowensis

Echinotriton raffaellii

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